About AJVR

American Journal of Veterinary Research The American Veterinary Medical Association
AJVR Staff  
Dr. Kurt J. Matushek,
Dr. Craig A. Smith,
Associate Editor
Dr. Helen L. Simons,
Associate Editor
Dr. Gussie J. Tessier,
Associate Editor
Dr. Alexandra L. Winter,
Assistant Editor
Dr. Rosemarie Seymour,
Assistant Editor
Dr. Christopher R. Byron,
Assistant Editor
Dr. Roxanne Pillars,
Assistant Editor
Dr. Bruce G. McLaughlin,
Adjunct Editor
Administrative Staff
Merri B. Benney,
Senior Assistant
to the Editor-in-Chief
Margaret M. Cafarella,
AJVR Secretary
Susan C. Kahler,
Senior News Editor
R. Scott Nolen,
Senior News Reporter
Katie Burns,
Senior News Reporter
Greg Cima,
News Reporter
Malinda Larkin,
News Reporter
Anita Suresh,
News Assistant Reporter
Graphic Design

Cheryl A. Atkins,
Design Services Manager

Justin D. Murray,

Graphic Designer 

Michele L. Morin,
Production Designer

​Diane A. Fagen,


Danielle Hildebrandt,
Library Technical Assistant

Nicholas T. De Luca,
Managing Editor
Justin D. DeAre,
Publications Technology Specialist
Linda G. Blatchford,
Online Technical Assistant
William A. Mazur,
Production Coordinator I
Victor L. Schultz,
Senior Copy Editor
Tram T. Le,
Copy Editor
Nicholas J. Roman,
Copy Editor
Lindy J. Paczak,
Copy Editor

Christina M. Banks,
Display Advertising Associate

Ian W. Tancun,
Advertising and Subscription
Fulfillment Associate 

AVMA Officers Business Information
Dr. Clark K. Fobian,
Dr. Theodore J. Cohn,
Dr. Walter R. Threlfall,
Vice President
Dr. Barbara A. Schmidt,
Dr. Douglas G. Aspros,
Immediate Past President
Dr. Thomas F. Meyer,
Executive Board Chair
Dr. V. Hugh "Chip" Price,
Executive Board Vice Chair
Dr. W. Ron DeHaven,
Executive Vice President
Dr. Elizabeth A. Curry-Galvin,
Assistant Executive Vice President
Editorial Office
1931 N Meacham Rd, Suite 100
Schaumburg, IL 60173-4360
Phone: 847-925-8070, ext 6752
Fax: 847-925-9329
E-mail: AJVR@avma.org

Web site: www.avma.org/journals

The American Journal of Veterinary Research (ISSN 0002-9645) is published monthly at 1931 N Meacham Rd, Suite 100, Schaumburg, IL 60173-4360. Periodicals postage paid at Schaumburg, Ill, and at additional mailing offices.

Opinions and statements expressed in the AJVR reflect the views of the contributors and do not represent the official policy of the American Veterinary Medical Association, unless so stated. Placement of an advertisement does not represent the AVMA's endorsement of the product or service.

Subscription price for nonmembers in the United States and possessions, $255 per year; for nonmembers in foreign countries, $265 per year. Single copies are $40 (United States and possessions) and $45 (foreign countries). For subscriptions and customer service, contact the AJVR by mail (AVMA M&FS Division, 1931 N Meacham Rd, Suite 100, Schaumburg, IL 60173-4360), phone (800-248-2862, ext 6639/6640), or e-mail (subscriptions@avma.org). For AVMA members, address changes can be made online at www.avma.org, by clicking on Member Center. Postmaster: send address changes to AJVR Address Changes, 1931 N Meacham Rd, Suite 100, Schaumburg, IL 60173-4360.

Material printed in the AJVR is covered by copyright. All rights reserved. No portion may be republished, reproduced (including photocopying), displayed, or transmitted without prior written permission from the publisher. Address permission requests to Diane A. Fagen, Permissions Coordinator (phone, 847-925-8070, ext 6770; fax, 847-925-9329; e-mail, dfagen@avma.org).

For author inquiries, contact Cindy Eyler at The Sheridan Press (phone, 800-352-2210, ext 8008; fax, 717-633-8929; e-mail, cindy.eyler@sheridan.com). For commercial inquiries, contact Beth Ann Rocheleau at Rockwater Inc (phone, 803-359-4578; fax, 803-753-9430; e-mail, brocheleau@ rockwaterinc.com).

Display Advertising  
For information on display advertising, contact Mark E. Reynolds at Media Communications Inc (phone, 913-541-9995; e-mail, mreynolds@avma.org).

AVMA District Directors
Dr. John H. de Jong
District I
Dr. Mark P. Helfat
District II
Dr. Michael E. Newman
District III
Dr. Larry G. Dee
District IV
Dr. Gary S. Brown
District V
Dr. Chester L. Rawson
District VI
Dr. John A. Howe
District VII
Dr. V. Hugh "Chip" Price
District VIII
Dr. Michael L. Whitehair
District IX
Dr. H. Theodore Trimmer
District X
Dr. Thomas F. Meyer
District XI
AVMA Information
The American Journal of Veterinary Research is owned and published by the American Veterinary Medical Association.

AVMA Offices
1931 N Meacham Rd, Suite 100
Schaumburg, IL 60173-4360
Phone: 847-925-8070 or
Fax: 847-925-1329

1910 Sunderland Place, NW
Washington, DC 20036-1642
Phone: 202-789-0007 or
Fax: 202-842-4360

AVMA Partners
AVMA Group Health
and Life Insurance Trust
3501 Frontage Rd
Tampa, FL 33607
Phone: 800-621-6360

PO Box 1629
Chicago, IL 60690-
Phone: 800-228-7548

American Veterinary Medical Foundation
1931 N Meacham Rd, Suite 100
Schaumburg, IL 60173-4360
Phone: 800-248-2862,
ext 6689